Stuart Lancaster has recalled Danny Care to the England squad for the tour of South Africa.
The Harlequins scrum half missed the Six Nations campaign after he was charged with drink driving.
James Haskell and Ugo Monye are also included after a spell out of the England fold, while Chris Robshaw is named as captain.
Hooker Tom Youngs joins brother Ben in the squad while there are also spaces for uncapped youngsters Christian Wade and Joe Launchbury in the 42-man party.
England will play three Tests against the Springboks throughout the course of June.
Full squad:
Forwards (23)
Mouritz Botha (Saracens), Dan Cole (Leicester Tigers), Alex Corbisiero (London Irish), Paul Doran Jones, Phil Dowson (both Northampton Saints), Carl Fearns (Bath Rugby), Joe Gray (Harlequins), Dylan Hartley (Northampton Saints), James Haskell (Otago Highlanders), Tom Johnson (Exeter Chiefs) Graham Kitchener (Leicester Tigers), Joe Launchbury (London Wasps), Joe Marler (Harlequins), Lee Mears (Bath Rugby), Ben Morgan (Scarlets), Matt Mullan (Worcester Warriors), Tom Palmer (Stade Francais), Geoff Parling (Leicester Tigers), Chris Robshaw (capt, Harlequins), George Robson (Harlequins), Matt Stevens (Saracens), Thomas Waldrom, Tom Youngs (both Leicester Tigers)
Backs (19)
Anthony Allen (Leicester Tigers), Chris Ashton (Northampton Saints), Brad Barritt (Saracens), Mike Brown, Danny Care (both Harlequins), Lee Dickson (Northampton Saints), Owen Farrell (Saracens), Toby Flood (Leicester Tigers), Ben Foden (Northampton Saints), Alex Goode (Saracens), Charlie Hodgson(Saracens), Jonathan Joseph (London Irish), George Lowe (Harlequins), Ugo Monye(Harlequins)
David Strettle (Saracens), Manu Tuilagi (Leicester Tigers), Jordan Turner-Hall (Harlequins), Christian Wade (London Wasps), Ben Youngs (Leicester Tigers)
