It had been suggested that the Chelsea defender would be omitted from the team in favour of Everton's Leighton Baines, but Hodgson has revealed that the 32-year-old will start at Wembley.
"Everyone has trained today, so we are all set for tomorrow. Ashley Cole will start tomorrow, I think it is important. He will start the game, but then I will see what I have to do with Leighton Baines, they are both exceptional players," he told reporters at a press conference.
Hodgson also confirmed that he will play his strongest available side as England attempt to beat Brazil for the first time in 23 years.